Heidi Harley
Professor, Psychology
B.A., University of Colorado at Boulder;
M.A., Ph.D., University of Hawaii at Manoa
Professor Harley is a comparative cognitive psychologist. Her research focuses on how the bottlenose dolphin represents its world. She has investigated questions concerning echolocation, dolphin whistles, spatial memory, rhythm processing, and imitation. Her courses include Introduction to Psychology, Cognitive Psychology, Comparative Cognition, Introduction to Environmental Studies, and Animal Language Research.
Recent Courses
Laboratory in Comparative Cognition
Cognitive Psychology
Psychology Senior Thesis Tutorial
Cognitive Psychology
Psychology Senior Thesis Tutorial
Selected Publications
Roitblat, H.L. & Harley, H.E. (1988). Rat spatial delayed matching-to-sample performance: Acquisition and retention. Journal of Experimental Psychology: Animal Behavior Processes, 14, 71-82.
Roitblat, H.L., Harley, H.E., & Helweg, D.A. (1993). Cognitive processing in artificial language research. In H.L. Roitblat, L.M. Herman, & P.E. Nachtigall (Eds.) Language and communication: Comparative perspectives (pp. 1-23). Hillsdale, NJ: Erlbaum.
Harley, H.E., Xitco, M.J., & Roitblat, H.L. (1995). Echolocation, cognition, and the dolphin's world. In R. Kastelein, J. Thomas, & P. Nachtigall (Eds.), Sensory Systems in Aquatic Mammals (pp. 515-528). Woerden, The Netherlands: De Spill.
Roitblat, H.L., Helweg, D.A., & Harley, H.E. (1995). Echolocation and imagery. In R. Kastelein, J. Thomas, & P. Nachtigall (Eds.), Sensory Systems in Aquatic Mammals (pp. 171-182). Woerden, The Netherlands: De Spill.
Harley, H.E., Roitblat, H.L. & Nachtigall, P.E. (1996). Object representation in the bottlenose dolphin (Tursiops truncatus): Integration of visual and echoic information. Journal of Experimental Psychology: Animal Behavior Processes, 22(2), 164-174 .
Harley, H.E., Putman, E.A., & Roitblat, H.L. (2003). Bottlenose dolphins perceive object features through echolocation. Nature, 424, 667-669.
Masters, M. & Harley, H.E. Introduction: Performance and cognition in echolocating mammals. (2004). In J. Thomas, C. Moss, & M. Vater (Eds.), Echolocation in Bats and Dolphins (pp. 249-259). Chicago: University of Chicago Press.
Delong, C.M., Au, W.W.L., Lemonds, D., Harley, H.E., & Roitblat, H.L. (2006). Acoustic features of objects matched by an echolocating bottlenose dolphin. Journal of the Acoustical Society of America, 119(3), 1867-1879.
Fellner, W., Bauer, G.B., & Harley, H.E. (2006). Cognitive implications of synchrony in dolphins: A review. Aquatic Mammals, 32(4), 511-516.
DeLong, C.M., Au, W.W.L., Harley, H.E., Roitblat, H.L., & Pytka, L. (2007). Human listeners provide insights into echo features used by dolphins to discriminate among objects. Journal of Comparative Psychology, 121(3), 306-319.
Harley, H.E. (2008). Whistle discrimination and categorization by the bottlenose dolphin (Tursiops truncatus): A review of the signature whistle framework and a perceptual test. Behavioural Processes, 77, 243-268.
Harley, H.E. & DeLong, C.M. (2008). Echoic object representation by the bottlenose dolphin. Comparative Cognition and Behavior Reviews, 3, 46-65.
Roitblat, H.L., Harley, H.E., & Helweg, D.A. (1993). Cognitive processing in artificial language research. In H.L. Roitblat, L.M. Herman, & P.E. Nachtigall (Eds.) Language and communication: Comparative perspectives (pp. 1-23). Hillsdale, NJ: Erlbaum.
Harley, H.E., Xitco, M.J., & Roitblat, H.L. (1995). Echolocation, cognition, and the dolphin's world. In R. Kastelein, J. Thomas, & P. Nachtigall (Eds.), Sensory Systems in Aquatic Mammals (pp. 515-528). Woerden, The Netherlands: De Spill.
Roitblat, H.L., Helweg, D.A., & Harley, H.E. (1995). Echolocation and imagery. In R. Kastelein, J. Thomas, & P. Nachtigall (Eds.), Sensory Systems in Aquatic Mammals (pp. 171-182). Woerden, The Netherlands: De Spill.
Harley, H.E., Roitblat, H.L. & Nachtigall, P.E. (1996). Object representation in the bottlenose dolphin (Tursiops truncatus): Integration of visual and echoic information. Journal of Experimental Psychology: Animal Behavior Processes, 22(2), 164-174 .
Harley, H.E., Putman, E.A., & Roitblat, H.L. (2003). Bottlenose dolphins perceive object features through echolocation. Nature, 424, 667-669.
Masters, M. & Harley, H.E. Introduction: Performance and cognition in echolocating mammals. (2004). In J. Thomas, C. Moss, & M. Vater (Eds.), Echolocation in Bats and Dolphins (pp. 249-259). Chicago: University of Chicago Press.
Delong, C.M., Au, W.W.L., Lemonds, D., Harley, H.E., & Roitblat, H.L. (2006). Acoustic features of objects matched by an echolocating bottlenose dolphin. Journal of the Acoustical Society of America, 119(3), 1867-1879.
Fellner, W., Bauer, G.B., & Harley, H.E. (2006). Cognitive implications of synchrony in dolphins: A review. Aquatic Mammals, 32(4), 511-516.
DeLong, C.M., Au, W.W.L., Harley, H.E., Roitblat, H.L., & Pytka, L. (2007). Human listeners provide insights into echo features used by dolphins to discriminate among objects. Journal of Comparative Psychology, 121(3), 306-319.
Harley, H.E. (2008). Whistle discrimination and categorization by the bottlenose dolphin (Tursiops truncatus): A review of the signature whistle framework and a perceptual test. Behavioural Processes, 77, 243-268.
Harley, H.E. & DeLong, C.M. (2008). Echoic object representation by the bottlenose dolphin. Comparative Cognition and Behavior Reviews, 3, 46-65.
